David Soto

Research

Group Leader. Ikerbasque Research Professor.

He leads the Consciousness Group at BCBL - Basque Center on Cognition Brain and Language.

Keywords: (Un)conscious information processing, metacognition, subjective experience, decoded neurofeedback, brain imaging, psychophysics.

David received his PhD in Experimental Psychology (2004) from the University of Santiago de Compostela. He then moved to the UK to join the Behavioural Brain Sciences Center at the University of Birmingham, where he was awarded a Postdoctoral Research Fellowship by the British Academy for the Humanities. In 2008, he established his own lab at the Division of Brain Sciences, Imperial College London, where he served as Assistant and later Associate Professor until 2016. During this period, he received the Medical Research Council's New Investigator Award and held visiting scholar positions at Harvard Medical School and the Neuroimaging Research Center at the University of Texas at Austin. In 2016, David returned to Spain to launch a new lab at the BCBL. His current research focuses on the distinction between conscious and non-conscious processes involved in perceptual selection, learning and memory, language, and metacognition. His team uses a broad range of behavioral, neuroimaging, and computational methods in both typical and atypical populations. The ultimate goal is to develop new protocols—for example, involving decoded neurofeedback—to manipulate underlying brain mechanisms and drive meaningful changes in cognition, behavior, and subjective experience.

Research group: Consciousness

d.soto@bcbl.eu

+34 943 309 300 Ext.: 211

2025

Mei, N., & Soto, D. (2025). Brain Representation in Conscious and Unconscious Vision. Journal of Cognition, 8(1). Doi:10.5334/joc.443

Olivers, C.N.L., Roelfsema, P.R., Soto, D., & Woodman, G.F. (2025). How Attention and Working Memory Married and Divorced, With Journal of Experimental Psychology: Human Perception and Performance as the Witness. Journal of Experimental Psychology Human Perception and Performance, 51(6), 689-692. Doi:10.1037/xhp0001267

Yan, K., Wokke, M.E., Wierzchoń, M., Wheatley, T., Weisberg, J., Webb, T., Watzl, S., Vibell, J., Tse, P.U., Tong, F., Taschereau-Dumouchel, V., Stein, T., Spiers, H., Soto, D., Solovey, G., Smith, B.C., Slagter, H.A., Silvanto, J., Shepherd, J., Seitz, A.R., Scott, R.B., Schwarzkopf, D.S., Schurger, A., Schiller, D., Roskies, A., Ringach, D.L., Rees, G., Rahnev, D., Rademaker, R.L., Phillips, I., Peters, M.A.K., Pestilli, F., Pereplyotchik, D., Owen, A.M., Or, C.C.F., Odegaard, B., Mylopoulos, M., Morales, J., Mok, R.M., Mobbs, D., Mitra, P.P., Miller, K.D., Michel, M., Metcalfe, J., Martinez-Trujillo, J., Machizawa, M.G., Liu, K.Y., Lin, Y.T., Lieberman, M.D., Libedinsky, C., Lee, A.L.F., LeDoux, J., Lau, H., Lanfranco, R.C., Kwok, S.C., Kvamme, T.L., Kording, K., Konstantinou, N., Knapen, T., Kentridge, R., Hung, S.M., Hsieh, P.J., Heyes, C., Herzog, M.H., Hassin, R.R., Hanson, J.R., Gottlieb, J., Goodale, M.A., Garfinkel, S., Frith, C.D., Frankish, K., Fleming, S.M., Dux, P.E., Doerig, A., Dijkstra, N., Dennett, D.C., De Brigard, F., de Gelder, B., Cushing, C.A., Cumming, S., Christoff, K., Corlett, P.R., Cohen, M.A., Cleeremans, A., Cherkaoui, M., Charest, I., Chang, D.H.F., Carter, O., Carruthers, P., Carrasco, M., Carmel, D., Buonomano, D., Browning, J., Bisley, J.W., Bengio, Y., Bekinschtein, T.A., Baxter, M.G., Arnold, D.H., Klincewicz, M., & Cheng, T. (2025). What makes a theory of consciousness unscientific?. Nature Neuroscience, 28(4), 689-693. Doi:10.1038/s41593-025-01881-x

2024

Soto, D., Salazar, A., Elosegi, P., Walter, A., Mei, N., Rodriguez, E., Petrollini, V., & Vicente, A. (2024). A novel image database for social concepts reveals preference biases in autistic spectrum in adults and children. Psychonomic Bulletin and Review, 31(4), 1690-1703. Doi:10.3758/s13423-023-02443-7

Elosegi, P., Mei, N., & Soto, D. (2024). Characterising the Role of Awareness in Ensemble Perception. Journal of Experimental Psychology Human Perception and Performance, 50(7), 706-722. Doi:10.1037/xhp0001201

Margolles, P., Elosegi, P., Mei, N., & Soto, D. (2024). Unconscious manipulation of conceptual representations with decoded neurofeedback impacts search behaviour. Journal of Neuroscience, 44(2). Doi:10.1523/JNEUROSCI.1235-23.2023

Alcalá-López, D., Mei, N., Margolles, P., & Soto, D. (2024). Brain-wide representation of social knowledge. Social Cognitive and Affective Neuroscience, 19(1). Doi:10.1093/scan/nsae032

Elosegi, P., Rahnev, D., & Soto, D. (2024). Think twice: Re-assessing confidence improves visual metacognition. Attention Perception and Psychophysics, 86(2), 373-380. Doi:10.3758/s13414-023-02823-0

Margolles, P., & Soto, D. (2024). Enhanced generalization and specialization of brain representations of semantic knowledge in healthy aging. Neuropsychologia, 204. Doi:10.1016/j.neuropsychologia.2024.108999

2023

Mei, N., Rahnev, D., & Soto, D. (2023). Using serial dependence to predict confidence across observers and cognitive domains. Psychonomic Bulletin and Review, 30(4), 1596-1608. Doi:10.3758/s13423-023-02261-x

Kartushina, N., Soto, D., & Martin, C. (2023). Metacognition in Second Language Speech Perception and Production. Language Learning, 73(2), 508-542. Doi:10.1111/lang.12549

Cudeiro, J., Soto, D., & Gutiérrez, E. (2023). Heat exposure following encoding can interfere with subsequent recognition memory. Scientific Reports, 13(1). Doi:10.1038/s41598-023-38248-w

Mazor, M., Brown, S., Ciaunica, A., Demertzi, A., Fahrenfort, J., Faivre, N., Francken, J.C., Lamy, D., Lenggenhager, B., Moutoussis, M., Nizzi, M.C., Salomon, R., Soto, D., Stein, T., & Lubianiker, N. (2023). The Scientific Study of Consciousness Cannot and Should Not Be Morally Neutral. Perspectives on Psychological Science, 18(3), 535-543. Doi:10.1177/17456916221110222

2022

Taouki, I., Lallier, M., & Soto, D. (2022). The role of metacognition in monitoring performance and regulating learning in early readers. Metacognition and Learning, 17(3), 921-948. Doi:10.1007/s11409-022-09292-0

Mei, N., Santana, R., & Soto, D. (2022). Informative neural representations of unseen contents during higher-order processing in human brains and deep artificial networks. Nature Human Behaviour, 6(5), 720-731. Doi:10.1038/s41562-021-01274-7

Cheng, P., Grover, S., Wen, W., Sankaranarayanan, S., Davies, S., Fragetta, J., Soto, D., & Reinhart, R.M.G. (2022). Dissociable rhythmic mechanisms enhance memory for conscious and nonconscious perceptual contents. Proceedings of the National Academy of Sciences of the United States of America, 119(44). Doi:10.1073/pnas.2211147119

Rahnev, D., Balsdon, T., Charles, L., de Gardelle, V., Denison, R., Desender, K., Faivre, N., Filevich, E., Fleming, S.M., Jehee, J., Lau, H., Lee, A.L.F., Locke, S.M., Mamassian, P., Odegaard, B., Peters, M., Reyes, G., Rouault, M., Sackur, J., Samaha, J., Sergent, C., Sherman, M.T., Siedlecka, M., Soto, D., Vlassova, A., & Zylberberg, A. (2022). Consensus Goals in the Field of Visual Metacognition. Perspectives on Psychological Science, 17(6), 1746-1765. Doi:10.1177/17456916221075615

Güldener, L., Jüllig, A., Soto, D., & Pollmann, S. (2022). Frontopolar activity carries feature information of novel stimuli during unconscious reweighting of selective attention. Cortex, 153, 146-165. Doi:10.1016/j.cortex.2022.03.024

2021

Sheikh, U.A., Carreiras, M., & Soto, D. (2021). Neurocognitive mechanisms supporting the generalization of concepts across languages. Neuropsychologia, 153. Doi:10.1016/j.neuropsychologia.2020.107740

Güldener, L., Jüllig, A., Soto, D., & Pollmann, S. (2021). Feature-Based Attentional Weighting and Re-weighting in the Absence of Visual Awareness. Frontiers in Human Neuroscience, 15. Doi:10.3389/fnhum.2021.610347

2020

Ordin, M., Polyanskaya, L., Soto, D., & Molinaro, N. (2020). Electrophysiology of statistical learning: Exploring the online learning process and offline learning product. European Journal of Neuroscience, 51(9), 2008-2022. Doi:10.1111/ejn.14657

Ordin, M., Polyanskaya, L., & Soto, D. (2020). Neural bases of learning and recognition of statistical regularities. Annals of the New York Academy of Sciences, 1467(1), 60-76. Doi:10.1111/nyas.14299

Ordin, M., Polyanskaya, L., & Soto, D. (2020). Metacognitive processing in language learning tasks is affected by bilingualism. Journal of Experimental Psychology Learning Memory and Cognition, 46(3), 529-538. Doi:10.1037/xlm0000739

Mei, N., Rankine, S., Olafsson, E., & Soto, D. (2020). Similar history biases for distinct prospective decisions of self-performance. Scientific Reports, 10(1). Doi:10.1038/s41598-020-62719-z

Soto, D., Sheikh, U.A., Mei, N., & Santana, R. (2020). Decoding and encoding models reveal the role of mental simulation in the brain representation of meaning. Royal Society Open Science, 7(5). Doi:10.1098/rsos.192043

Rahnev, D., Desender, K., Lee, A.L.F., Adler, W.T., Aguilar-Lleyda, D., Akdoğan, B., Arbuzova, P., Atlas, L.Y., Balcı, F., Bang, J.W., Bègue, I., Birney, D.P., Brady, T.F., Calder-Travis, J., Chetverikov, A., Clark, T.K., Davranche, K., Denison, R.N., Dildine, T.C., Double, K.S., Duyan, Y.A., Faivre, N., Fallow, K., Filevich, E., Gajdos, T., Gallagher, R.M., de Gardelle, V., Gherman, S., Haddara, N., Hainguerlot, M., Hsu, T.Y., Hu, X., Iturrate, I., Jaquiery, M., Kantner, J., Koculak, M., Konishi, M., Koß, C., Kvam, P.D., Kwok, S.C., Lebreton, M., Lempert, K.M., Ming Lo, C., Luo, L., Maniscalco, B., Martin, A., Massoni, S., Matthews, J., Mazancieux, A., Merfeld, D.M., O’Hora, D., Palser, E.R., Paulewicz, B., Pereira, M., Peters, C., Philiastides, M.G., Pfuhl, G., Prieto, F., Rausch, M., Recht, S., Reyes, G., Rouault, M., Sackur, J., Sadeghi, S., Samaha, J., Seow, T.X.F., Shekhar, M., Sherman, M.T., Siedlecka, M., Skóra, Z., Song, C., Soto, D., Sun, S., van Boxtel, J.J.A., Wang, S., Weidemann, C.T., Weindel, G., Wierzchoń, M., Xu, X., Ye, Q., Yeon, J., Zou, F., & Zylberberg, A. (2020). The Confidence Database. Nature Human Behaviour, 4(3), 317-325. Doi:10.1038/s41562-019-0813-1

Li, K., Bentley, P., Nair, A., Halse, O., Barker, G., Russell, C., Soto, D., & Malhotra, P.A. (2020). Reward sensitivity predicts dopaminergic response in spatial neglect. Cortex, 122, 213-224. Doi:10.1016/j.cortex.2018.09.002

2019

Sheikh, U.A., Carreiras, M., & Soto, D. (2019). Decoding the meaning of unconsciously processed words using fMRI-based MVPA. Neuroimage, 191, 430-440. Doi:10.1016/j.neuroimage.2019.02.010

Bourbon-Teles, J., & Soto, D. (2019). Assessing the Role of the Left Dorsal Frontal Cortex in Working Memory Guidance: Attentional or Mnemonic? A Neurostimulation Study. Neuroscience, 411, 140-149. Doi:10.1016/j.neuroscience.2019.04.049

Soto, D., Sheikh, U.A., & Rosenthal, C.R. (2019). A Novel Framework for Unconscious Processing. Trends in Cognitive Sciences, 23(5), 372-376. Doi:10.1016/j.tics.2019.03.002

2018

Soto, D., Theodoraki, M., & Paz-Alonso, P.M. (2018). How the human brain introspects about one's own episodes of cognitive control. Cortex, 107, 110-120. Doi:10.1016/j.cortex.2017.10.016

Rosenthal, C.R., Mallik, I., Caballero-Gaudes, C., Sereno, M.I., & Soto, D. (2018). Learning of goal-relevant and -irrelevant complex visual sequences in human V1. Neuroimage, 179, 215-224. Doi:10.1016/j.neuroimage.2018.06.023

Rinne, P., Hassan, M., Fernandes, C., Han, E., Hennessy, E., Waldman, A., Sharma, P., Soto, D., Leech, R., Malhotra, P.A., & Bentley, P. (2018). Motor dexterity and strength depend upon integrity of the attention-control system. Proceedings of the National Academy of Sciences of the United States of America, 115(3), E536-E545. Doi:10.1073/pnas.1715617115

2017

Wedmore, F., Musil, S., & Soto, D. (2017). How ageing changes the mnemonic bias of visual behaviour. Visual Cognition, 25(1-3), 51-59. Doi:10.1080/13506285.2017.1330789

Soto, D. (2017). How Do We Keep Information ‘Online’?. Trends in Cognitive Sciences, 21(2), 63-65. Doi:10.1016/j.tics.2016.12.009

2016

Rosenthal, C.R., & Soto, D. (2016). The Anatomy of Non-conscious Recognition Memory. Trends in Neurosciences, 39(11), 707-711. Doi:10.1016/j.tins.2016.09.005

Li, K., Russell, C., Balaji, N., Saleh, Y., Soto, D., & Malhotra, P.A. (2016). The effects of motivational reward on the pathological attentional blink following right hemisphere stroke. Neuropsychologia, 92, 190-196. Doi:10.1016/j.neuropsychologia.2016.03.037

Rosenthal, C.R., Andrews, S.K., Antoniades, C.A., Kennard, C., & Soto, D. (2016). Learning and recognition of a non-conscious sequence of events in human primary visual cortex. Current Biology, 26(6), 834-841. Doi:10.1016/j.cub.2016.01.040

Soto, D., & Silvanto, J. (2016). Is conscious awareness needed for all working memory processes?. Neuroscience of Consciousness, 1. Doi:10.1093/nc/niw009

2015

Greene, C.M., Kennedy, K., & Soto, D. (2015). Dynamic states in working memory modulate guidance of visual attention: Evidence from an n-back paradigm. Visual Cognition, 23(5), 546-560. Doi:10.1080/13506285.2015.1052033

Nigmatullina, Y., Arshad, Q., Wu, K., Seemungal, B.M., Bronstein, A.M., & Soto, D. (2015). How imagery changes self-motion perception. Neuroscience, 291, 46-52. Doi:10.1016/j.neuroscience.2015.01.021

Jachs, B., Blanco, M.J., Grantham-Hill, S., & Soto, D. (2015). On the independence of visual awareness and metacognition: A signal detection theoretic analysis. Journal of Experimental Psychology Human Perception and Performance, 41(2), 269-276. Doi:10.1037/xhp0000026

Greene, C.M., Vidaki, K., & Soto, D. (2015). Fractionating the neural substrates of incidental recognition memory. Learning and Memory, 22(1), 24-30. Doi:10.1101/lm.036327.114

2014

Kiyonaga, A., Korb, F.M., Lucas, J., Soto, D., & Egner, T. (2014). Dissociable causal roles for left and right parietal cortex in controlling attentional biases from the contents of working memory. Neuroimage, 100, 200-205. Doi:10.1016/j.neuroimage.2014.06.019

De Bourbon-Teles, J., Bentley, P., Koshino, S., Shah, K., Dutta, A., Malhotra, P., Egner, T., Husain, M., & Soto, D. (2014). Thalamic control of human attention driven by memory and learning. Current Biology, 24(9), 993-999. Doi:10.1016/j.cub.2014.03.024

Dutta, A., Shah, K., Silvanto, J., & Soto, D. (2014). Neural basis of non-conscious visual working memory. Neuroimage, 91, 336-343. Doi:10.1016/j.neuroimage.2014.01.016

Soto, D., Rotshtein, P., & Kanai, R. (2014). Parietal structure and function explain human variation in working memory biases of visual attention. Neuroimage, 89, 289-296. Doi:10.1016/j.neuroimage.2013.11.036

Pan, Y., Lin, B., Zhao, Y., & Soto, D. (2014). Working memory biasing of visual perception without awareness. Attention Perception and Psychophysics, 76(7), 2051-2062. Doi:10.3758/s13414-013-0566-2

Greene, C.M., Flannery, O., & Soto, D. (2014). Distinct parietal sites mediate the influences of mood, arousal, and their interaction on human recognition memory. Cognitive Affective and Behavioral Neuroscience, 14(4), 1327-1339. Doi:10.3758/s13415-014-0266-y

Greene, C.M., & Soto, D. (2014). Functional connectivity between ventral and dorsal frontoparietal networks underlies stimulus-driven and working memory-driven sources of visual distraction. Neuroimage, 84, 290-298. Doi:10.1016/j.neuroimage.2013.08.060

Soto, D., & Silvanto, J. (2014). Reappraising the relationship between working memory and conscious awareness. Trends in Cognitive Sciences, 18(10), 520-525. Doi:10.1016/j.tics.2014.06.005

2013

Rinne, P., Hassan, M., Goniotakis, D., Chohan, K., Sharma, P., Langdon, D., Soto, D., & Bentley, P. (2013). Triple dissociation of attention networks in stroke according to lesion location. Neurology, 81(9), 812-820. Doi:10.1212/WNL.0b013e3182a2ca34

Bona, S., Cattaneo, Z., Vecchi, T., Soto, D., & Silvanto, J. (2013). Metacognition of visual short-term memory: Dissociation between objective and subjective components of VSTM. Frontiers in Psychology, 4(FEB). Doi:10.3389/fpsyg.2013.00062

Malhotra, P.A., Soto, D., Li, K., & Russell, C. (2013). Reward modulates spatial neglect. Journal of Neurology Neurosurgery and Psychiatry, 84(4), 366-369. Doi:10.1136/jnnp-2012-303169

2012

Soto, D., Greene, C.M., Kiyonaga, A., Rosenthal, C.R., & Egner, T. (2012). A parieto-medial temporal pathway for the strategic control over working memory biases in human visual attention. Journal of Neuroscience, 32(49), 17563-17571. Doi:10.1523/JNEUROSCI.2647-12.2012

Greene, C.M., & Soto, D. (2012). Neural repetition effects in the medial temporal lobe complex are modulated by previous encoding experience. Plos One, 7(7). Doi:10.1371/journal.pone.0040870

Soto, D., Greene, C.M., Chaudhary, A., & Rotshtein, P. (2012). Competition in working memory reduces frontal guidance of visual selection. Cerebral Cortex, 22(5), 1159-1169. Doi:10.1093/cercor/bhr190

Balani, A.B., Soto, D., & Humphreys, G.W. (2012). Separating top-down and bottom-up cueing of attention from response inhibition in utilization behavior. Neurocase, 18(2), 98-111. Doi:10.1080/13554794.2011.568496

Soto, D., Llewelyn, D., & Silvanto, J. (2012). Distinct causal mechanisms of attentional guidance by working memory and repetition priming in early visual cortex. Journal of Neuroscience, 32(10), 3447-3452. Doi:10.1523/JNEUROSCI.6243-11.2012

Silvanto, J., & Soto, D. (2012). Causal evidence for subliminal percept-to-memory interference in early visual cortex. Neuroimage, 59(1), 840-845. Doi:10.1016/j.neuroimage.2011.07.062

Kiyonaga, A., Egner, T., & Soto, D. (2012). Cognitive control over working memory biases of selection. Psychonomic Bulletin and Review, 19(4), 639-646. Doi:10.3758/s13423-012-0253-7

Särkämö, T., & Soto, D. (2012). Music listening after stroke: Beneficial effects and potential neural mechanisms. Annals of the New York Academy of Sciences, 1252(1), 266-281. Doi:10.1111/j.1749-6632.2011.06405.x

Soto, D., Rotshtein, P., Hodsoll, J., Mevorach, C., & Humphreys, G.W. (2012). Common and distinct neural regions for the guidance of selection by visuoverbal information held in memory: Converging evidence from fMRI and rTMS. Human Brain Mapping, 33(1), 105-120. Doi:10.1002/hbm.21196

2011

Soto, D., Mannan, S.K., Malhotra, P., Rzeskiewicz, A., & Humphreys, G.W. (2011). Distinguishing non-spatial from spatial biases in visual selection: Neuropsychological evidence. Acta Psychologica, 137(2), 226-234. Doi:10.1016/j.actpsy.2010.08.010

Soto, D., Mok, A.Y.F., McRobbie, D., Quest, R., Waldman, A., & Rotshtein, P. (2011). Biasing visual selection: Functional neuroimaging of the interplay between spatial cueing and feature memory guidance. Neuropsychologia, 49(6), 1537-1543. Doi:10.1016/j.neuropsychologia.2010.11.035

Rotshtein, P., Soto, D., Grecucci, A., Geng, J.J., & Humphreys, G.W. (2011). The role of the pulvinar in resolving competition between memory and visual selection: A functional connectivity study. Neuropsychologia, 49(6), 1544-1552. Doi:10.1016/j.neuropsychologia.2010.12.002

Soto, D., Mäntylä, T., & Silvanto, J. (2011). Working memory without consciousness. Current Biology, 21(22), R912-R913. Doi:10.1016/j.cub.2011.09.049

2010

Balani, A.B., Soto, D., & Humphreys, G.W. (2010). Working memory and target-related distractor effects on visual search. Memory and Cognition, 38(8), 1058-1076. Doi:10.3758/MC.38.8.1058

Rosenthal, C.R., Kennard, C., & Soto, D. (2010). Visuospatial sequence learning without seeing. Plos One, 5(7). Doi:10.1371/journal.pone.0011906

Greene, C.M., Bahri, P., & Soto, D. (2010). Interplay between affect and arousal in recognition memory. Plos One, 5(7). Doi:10.1371/journal.pone.0011739

Mannan, S.K., Kennard, C., Potter, D., Pan, Y., & Soto, D. (2010). Early oculomotor capture by new onsets driven by the contents of working memory. Vision Research, 50(16), 1590-1597. Doi:10.1016/j.visres.2010.05.015

Pan, Y., & Soto, D. (2010). The modulation of perceptual selection by working memory is dependent on the focus of spatial attention. Vision Research, 50(15), 1437-1444. Doi:10.1016/j.visres.2009.10.016

Grecucci, A., Soto, D., Rumiati, R.I., Humphreys, G.W., & Rotshtein, P. (2010). The interrelations between verbal working memory and visual selection of emotional faces. Journal of Cognitive Neuroscience, 22(6), 1189-1200. Doi:10.1162/jocn.2009.21276

Soto, D., Wriglesworth, A., Bahrami-Balani, A., & Humphreys, G.W. (2010). Working Memory Enhances Visual Perception: Evidence From Signal Detection Analysis. Journal of Experimental Psychology Learning Memory and Cognition, 36(2), 441-456. Doi:10.1037/a0018686

2009

Balani, A.B., Soto, D., & Humphreys, G.W. (2009). Constraints on task-based control of behaviour following frontal lobe damage: A single-case study. Cognitive Neuropsychology, 26(7), 635-654. Doi:10.1080/02643291003633827

Kumar, S., Soto, D., & Humphreys, G.W. (2009). Electrophysiological evidence for attentional guidance by the contents of working memory. European Journal of Neuroscience, 30(2), 307-317. Doi:10.1111/j.1460-9568.2009.06805.x

Soto, D., & Humphreys, G.W. (2009). Automatic selection of irrelevant object features through working memory evidence for top-down attentional capture. Experimental Psychology, 56(3), 165-172. Doi:10.1027/1618-3169.56.3.165

Soto, D., Funes, M.J., Guzmán-García, A., Warbrick, T., Rotshtein, P., & Humphreys, G.W. (2009). Pleasant music overcomes the loss of awareness in patients with visual neglect. Proceedings of the National Academy of Sciences of the United States of America, 106(14), 6011-6016. Doi:10.1073/pnas.0811681106

Soto, D., Montoro, P.R., & Humphreys, G.W. (2009). Transcranial magnetic stimulation of the primary motor cortex modulates response interference in a flanker task. Neuroscience Letters, 451(3), 261-265. Doi:10.1016/j.neulet.2008.12.052

Blanco, M.J., & Soto, D. (2009). Unconscious perception of a flash can trigger line motion illusion. Experimental Brain Research, 192(4), 605-613. Doi:10.1007/s00221-008-1564-9

Soto, D., & Humphreys, G.W. (2009). Semantically induced distortions of visual awareness in a patient with Balint's syndrome. Cognition, 110(2), 237-241. Doi:10.1016/j.cognition.2008.08.006

Pan, Y., Xu, B., & Soto, D. (2009). Dimension-based working memory-driven capture of visual selection. Quarterly Journal of Experimental Psychology, 62(6), 1123-1131. Doi:10.1080/17470210802624353

Blanco, M.J., Leirós, L.I., & Soto, D. (2009). The inter-relations between spatial and object factors during sustained visual attention to dynamic objects. Psicologica, 30(2), 271-285.

Soto, D. (2009). Interplay between music, emotion and cognitive function in health and disease. Communicative and Integrative Biology, 2(6), 549-551. Doi:10.4161/cib.2.6.9656

2008

Soto, D., & Humphreys, G.W. (2008). Stressing the mind: The effect of cognitive load and articulatory suppression on attentional guidance from working memory. Perception and Psychophysics, 70(5), 924-934. Doi:10.3758/PP.70.5.924

Valle-Inclán, F., Blanco, M.J., Soto, D., & Leirós, L. (2008). A new method to assess eye dominance. Psicologica, 29(1), 55-64.

Soto, D., Hodsoll, J., Rotshtein, P., & Humphreys, G.W. (2008). Automatic guidance of attention from working memory. Trends in Cognitive Sciences, 12(9), 342-348. Doi:10.1016/j.tics.2008.05.007

2007

Soto, D., Humphreys, G.W., & Rotshtein, P. (2007). Dissociating the neural mechanisms of memory-based guidance of visual selection. Proceedings of the National Academy of Sciences of the United States of America, 104(43), 17186-17191. Doi:10.1073/pnas.0703706104

Soto, D., & Humphreys, G.W. (2007). Automatic Guidance of Visual Attention From Verbal Working Memory. Journal of Experimental Psychology Human Perception and Performance, 33(3), 730-737. Doi:10.1037/0096-1523.33.3.730

2006

Soto, D., Humphreys, G.W., & Heinke, D. (2006). Dividing the mind: The necessary role of the frontal lobes in separating memory from search. Neuropsychologia, 44(8), 1282-1289. Doi:10.1016/j.neuropsychologia.2006.01.029

Soto, D., & Humphreys, G.W. (2006). Seeing the content of the mind: Enhanced awareness through working memory in patients with visual extinction. Proceedings of the National Academy of Sciences of the United States of America, 103(12), 4789-4792. Doi:10.1073/pnas.0510718103

Soto, D., Humphreys, G.W., & Heinke, D. (2006). Working memory can guide pop-out search. Vision Research, 46(6-7), 1010-1018. Doi:10.1016/j.visres.2005.09.008

2005

Soto, D., Heinke, D., Humphreys, G.W., & Blanco, M.J. (2005). Early, involuntary top-down guidance of attention from working memory. Journal of Experimental Psychology Human Perception and Performance, 31(2), 248-261. Doi:10.1037/0096-1523.31.2.248

2004

Soto, D., & Blanco, M.J. (2004). Spatial attention and object-based attention: A comparison within a single task. Vision Research, 44(1), 69-81. Doi:10.1016/j.visres.2003.08.013

2003

Blanco, M.J., & Soto, D. (2003). Compensating small changes in luminance during performance of a visual sustained attention task. Travail Humain, 66(4), 329-346. Doi:10.3917/th.664.0329

2002

Blanco, M.J., & Soto, D. (2002). Effects of spatial attention on detection and identification of oriented lines.. Acta Psychologica, 109(2), 195-212. Doi:10.1016/S0001-6918(01)00057-9

2024

CODE: COnfident DEcisions

Program: MSCA Doctoral network 2022, MSCA-2022-DN_101119647
Funding agency: European Commission
09/2024- 08/2027
Funding: 251.971 €
PI: David Soto PhD Student: Francisca Campos Matias
A-EPIC: Assessing Awareness in Ensemble Perception through Optimized Psychophysics and Closed-Loop Brain Imaging

Program: Plan Nacional 2023, PID2023-149267NB-I00
Funding agency: Spanish Ministry
09/2024- 08/2028
Funding: 170.125 €
PI: David Soto Team member: Najemeddine Abdennour, Yining (Ivory) Cheng

2020

BrainMetaLoop: Using closed-loop brain imaging to understand and manipulate metacognitive functions

Program: Plan Nacional 2019, PID2019-105494GB-I00
Funding agency: Spanish Ministry
06/2020- 05/2024
Funding: 128.000 €
Pi: David Soto

2017

La interacción entre la memoria operativa y la atención visual a lo largo del ciclo vital

Program: PIBA 2017, PI_2017_1_0025
Funding agency: Basque Government
09/2017- 09/2019
Funding: 69.600 €
PI: David Soto

2016

METAWARE: Brain mechanisms for human working memory and metacognition across different states of awareness

Program: Plan Nacional 2016, PSI2016-76443-P
Funding agency: Spanish Ministry
12/2016- 12/2019
Funding: 48.000 €
PI: David Soto
Centro de Excelencia Severo Ochoa: BCBL

Program: SEV-2015-0490
Funding agency: Spanish Ministry
01/2016- 01/2019
Funding: 4.000.000 €
PI: Prof. Manuel Carreiras Guarantors: Arthur Samuel, David Soto, Clara D. Martin, Nicola Molinaro, Mari Cruz Rodriguez-Oroz, Marie Lallier, Kepa Paz-Alonso, Doug Davidson, Simona Mancini & Loretxu Bergouignan

In progress

A-EPIC-Assessing Awareness in Ensemble Perception through Optimized Psychophysics and Closed-Loop Brain Imaging University of the Basque Country, UPV/EHU (2025 - 2029)

The role of collaboration in decision making and metacognition in childhood University of the Basque Country, UPV/EHU (2024 - 2028)

Investigating visual conscious experiences using ensemble perception University of the Basque Country, UPV/EHU (2023 - 2027)

Aha-Moment through Prediction Error University of the Basque Country, UPV/EHU (2021 - 2025)

Defended

The Role of Perceptual Awareness in Ensemble Perception: Psychophysical and Neural Mechanisms University of the Basque Country, UPV/EHU (2021 - 2025)

Domain-general mechanisms and language recovery after stroke: a longitudinal study University of the Basque Country, UPV/EHU (2020 - 2025)

The dynamic nature of conceptual representations in the human brain: insights from aging and decoded neurofeedback University of the Basque Country, UPV/EHU (2018 - 2023)

Assessing the representation of seen and unseen contents in human brains and deep artificial networks University of the Basque Country, UPV/EHU (2018 - 2022)

The role of metacognitive monitoring in regulating learning in early readers University of the Basque Country, UPV/EHU (2017 - 2021)

The Neural Representation of Concepts in Bilinguals: An Evaluation of Factors Influencing Cross-language Overlap. Using fMRI-based Multivariate Pattern Analysis University of the Basque Country, UPV/EHU (2016 - 2023)

Magnocellular and parvocellular visual pathways' involvement in reading University of the Basque Country, UPV/EHU (2016 - 2022)

Attention-control deficits and their impact upon motor deficits in stroke Imperial College of London (2016 - 2016)

Neural substrates supporting the influence of working memory contents on visual attention Imperial College of London (2014 - 2014)